YIELD STABILITY ANALYSIS IN TARO (COLOCASIA ESCULENTA) AS INFLUENCED BY SOME CROP MANAGEMENT SYSTEMS

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ABSTRACT

The study was carried out to assess the yield stability of some taro (Colocasia esculenta (L.) Schott) genotypes commonly grown by farmers in Nigeria as influenced by some cropping system management practices (two locations in two years). The experiment was laid out as a split plot in a randomized complete block design with three replications. The six genotypes used were: NCe 001, NCe 002, NCe 003, NCe 004, NCe 005 and NCe 010. Data were collected on the growth characteristics and yield attributes. There was significant effect of crop management system on most of the growth and yield attributes measured. The genotypes varied significantly (p<0.001) in their performance for almost all the traits studied. Location had significant effect on taro yield with Ishiagu giving the best performance for growth and most of the measured yield on attributes. Genotypes had strong influence all the characters considered in this study with NCe 002 and NCe004 consistently giving the best performance in yield and few of the studied traits. This indicates that these genotypes show good adaptation to the soil and climatic conditions of these agro-ecosystems. AMMI showed high significance on the taro yieldEffects of genotype and environment were highly significant. The main effects (genotypes and environments) captured 62.9% of the total sum of squares (TSS) while the genotype by environment interaction (GEI) contained 8.2% of the TSS for population density. Under propagule the main effects (genotype and environment) captured 52.1% of the total sum of squares (TSS) while the GEI explained 17.2% of the total variation. For fertilizer rates, the main effects (genotypes and environments) captured 87.6% of the total sum of squares (TSS) while the GEI explained 4.5% of the TSS. The linear correlation (r) coefficients between the various attributes and the yield as affected by fertilizer rates showed that the correlation in number of secondary suckers and cormel number per plant was positively and significantly associated with yield (t/ha), r = 0.223** and r = 0.169*, respectively. However, plant height, number of leaves, leaf area, cormel length and cormel circumference were negatively but significantly associated with yield (t/ha). With regards to the linear correlation (r) coefficients between the various attributes and the yield per hectare as affected by propagule sizes, the cormel number per plant and cormel weight per plant were positively and significantly associated with yield (t/ha), r = 0.549** and r = 0.330*, respectively. However, plant height, number of leaves, number of secondary suckers and leaf area were negatively but significantly associated with yield (t/ha). In 2016, plant h==[-eight was the largest contributor to yield accounting for up to 18 % of the total yield (B = 0.421), while cormel number per plant was observed to be the largest contributor to yield accounting for up to 38 % of the total yield (B = 0.619; P<0.001) in 2017.





TABLE OF CONTENTS

Title Page                                                                                                                    i

Declaration                                                                                                                 ii

Certification                                                                                                               iii

Dedication                                                                                                                  iv

Acknowledgements                                                                                                    v

Table of Contents                                                                                                       vi

List of Tables                                                                                                              xi

List of Figures                                                                                                             xvi

Abstract                                                                                                                      xvii

 

CHAPTER 1: INTRODUCTION                                                                            1

CHAPTER 2: LITERATURE REVIEW                                                               7

2.1       Origin and Domestication of Taro                                                                  7

2.2       Taxonomical Description of Taro                                                                  8

2.3       Botany and Ecology of Taro                                                                          11

2.3.1    Classification of taro                                                                                      11

2.4       Taro Morphology and Anatomy                                                                     11

2.5       Growth Cycle and Developmental Stages of Taro                                         14

2.5.1    Establishment                                                                                                 15

2.5.2    Vegetative growth and corm initiation                                                           15

2.5.3    Corm bulking and maturation                                                                         16

2.6       Nutritional Content of Taro                                                                            16

2.7       Genetic Diversity Assessment of Taro                                                           18

2.8       Methods of Assessing Genetic Diversity                                                       20

2.8.1    Morphological markers                                                                                  20

2.8.2    Genetic markers                                                                                              21

2.9       Economic Importance and Uses of Taro                                                        23

2.10     Yield and Yield Attributes                                                                             25

2.11     Environmental Requirements for the Cultivation of Taro                             29

2.11.1  Water                                                                                                              29

2.11.2  Photoperiod and light intensity                                                                       30

2.11.3  Soils                                                                                                                31

2.12     Taro Cultivation                                                                                             31

2.13     Taro Planting                                                                                                  32

2.14     Planting Density                                                                                             32

2.15     Factors Influencing Taro Quality                                                                   33

2.15.1  Genetic factors                                                                                                34

2.15.2  Planting date                                                                                                   35

2.15.3  Temperature                                                                                                   35

2.15.4  Rainfall                                                                                                           37

2.16     Fertilizer Rate                                                                                                 38

 

CHAPTER 3: MATERIALS AND METHODS                                                    42

3.1       Experiment I: Genotypes by Planting Density Interaction in Some Taro     Genotypes                                                                                                       42

 

3.2       Experiment II: Effect of Different Fertilizer Rates on the Yield Stability

            of Some Taro Genotypes                                                                                45

3.3       Experiment III: Effects of Size of Propagules on Yield Stability of Some

            Taro Genotypes                                                                                              46

 

CHAPTER 4: RESULTS                                                                                         49

4.1       Soil Physicochemical Properties of Experimental Sites                                49

4.2       Agro-Meteorological Data of the Experimental Sites                                    49

4.3       Effects of Planting Density on Growth, Yield and Yield Stability of

            Some Taro Genotypes                                                                                    54

4.3.1    Effects of planting density on growth attributes                                            54

4.3.1.1. Effect of planting density on plant height (cm) of some taro genotypes

            at 8, 12, 16 and 20 WAP in two locations                                                      54

4.3.1.2.Effect of planting density on number of leaves of some taro genotypes

            at 8, 12, 16 and 20 WAP in two locations                                                      60

4.3.1.3 Effect of planting density on leaf area of some taro genotypes at 12,

16 and 20 WAP in two locations                                                                 64

4.3.1.4 Effect of planting density on number of secondary suckers of

            some taro genotypes at 12, 16 and 20 WAP in two locations                                  69

4.3.2    Effect of planting density on yield and yield associated traits                     72

4.3.2.1 Effect of planting density on taro yield                                                          72

4.3.2.2 Effect of planting density on cormel length                                                   74

4.3.2.3 Effect of planting density on cormel circumference                                      74

4.3.2.4 Effect of planting density on number of cormels per plant                            77

4.3.2.5 Effect of planting density on cormel weight per plant                                               as influenced by plant population density                                                      79

4.3.2.6 Effect of planting density on corm length                                                      82

4.3.2.7 Effect of planting density on corm circumference                                         84

4.3.2.8 Effect of planting density on corm weight per plant                                      84

4.4       Effects of Different Fertilizer Rates on Growth, Yield and Yield

            Stability of Some Taro Genotypes                                                                 86

4.4.1    Effects of fertilizer rates on growth attributes                                                86

4.4.1.1 Effect of fertilizer rate on plant height (cm) of some taro genotypes

            at 8, 12, 16 and 20 WAP in two locations                                                      86

4.4.1.2 Effect of fertilizer rate on number of leaves of some taro genotypes

            at 8, 12, 16 and 20 WAP in two locations                                                      92

4.4.1.3 Leaf area (cm2) at 8, 12, 16 and 20 WAP of some taro genotypes as

            affected by location and fertilizer rates.                                                         98

4.4.1.4 Number of secondary suckers at 8, 12, 16 and 20 WAP of some taro

genotypes as influenced by location and fertilizer rates.                               104

4.4.2    Effect of fertilizer rates on yield and its associated traits                              110

4.4.2.1 Effect of fertilizer rates on yield per hectare per tonne                                  110

4.4.2.2 Effect of fertilizer rates on corm circumference                                            112

4.4.2.3 Effect of fertilizer rates on corms length                                                        112

4.4.2.4 Effect of fertilizer rates on cormel length                                                      112

4.4.2.5 Effect of fertilizer rate on corm weight per plant                                           115

4.4.2.6 Effect of fertilizer rate on cormel circumference                                           115

4.4.2.7 Effect of fertilizer rate on number of cormels per plant                                 115

4.5       Effects of Propagule Size on Growth, Yield and Yield Stability of

            Some Taro Genotypes                                                                                    119

4.5.1    Effects of propagule size on growth attributes                                               119

4.5.1.1 Effect of propagule size on plant height (cm) of some taro genotypes

            at 8, 12, 16 and 20 WAP in two locations                                                      119

4.5.1.2 Effect of propagule size on number of leaves of some taro genotypes

at 8, 12 and 16 WAP in two locations                                                            125

4.5.1.3 Effect of propagule weight on leaf area of some taro genotypes at 8, 12,

16 and 20 WAP in two locations                                                                    128

4.5.1.4 Effect of propagule weight on number of secondary suckers of

some taro genotypes at 8, 12, 16 and 20 WAP in two locations                  133

4.5.2    Effect of propagule size on yield and yield-associated traits                         138

4.5.2.1 Effect of propagule size on yield per hectare                                                 138

4.5.2.2 Effect propagule size on corm circumference                                               138

4.5.2.3 Effect propagule size on corm length                                                             141

4.5.2.4 Effect propagule size on cormel circumference                                             141

4.5.2.5 Effect propagule size on cormel length                                                          141

4.5.2.6 Effect propagule size on number of cormels per plant                                   145

4.5.2.7 Effect propagule size on cormel weight per plant                                          147

4.5.2.8 Effect propagule size on corm weight per plant                                             147

4.6       Yield Stability Analyses                                                                                 150

4.6.1    Population density                                                                                          150

4.6.2    Propagule size                                                                                                 159

4.6.3    Fertilizer rates                                                                                                 168

4.7           Correlation Analysis                                                                                     176

4.7.1    Correlation analysis of the growth and yield attributes of taro as

            affected by the fertilizer rates in the first planting year (2016)                    176

4.7.2    Correlation analysis of the growth and yield attributes of taro as affected

            by the fertilizer rates in the second planting year (2017)                              178

4.7.3    Correlation analysis of the growth and yield attributes of taro as affected

            by the propagule size in the first planting year                                             180

4.7.4    Correlation analysis of the growth and yield attributes of taro as affected

            by the propagule size in the second planting year                                         182

4.8           Multiple Regression Studies                                                                         184

 

CHAPTER 5: DISCUSSION AND CONCLUSION                                             183

References

 

 

 

 

 

 

 

 

 

 

LIST OF TABLES


 4.1:     Soil physico-chemical characteristics of the study site for 2016

             and 2017 cropping season.                                                                            50

 4.2:     Agro-meteorogical data of the experimental sites                                          52

 4.3:     Agro-meteorogical data of the experimental sites 2017                                 53

 4.4:     Plant height (cm) of some taro genotypes at 8 WAP across two locations

            in two planting seasons                                                                                   56

 4.5:     Plant height (cm) of some taro genotypes at 12 WAP across two

            locations in two planting seasons                                                                   57

 4.6:     Plant height (cm) of some taro genotypes at 16 WAP across two

            locations in two planting seasons                                                                   58

 4.7:     Plant height (cm) of some taro genotypes at 20 WAP across two locations

            in two planting seasons                                                                                   59

 4.8:     Number of leaves of some taro genotypes at 12 WAP across two

            locations in two planting seasons                                                                   61

 4.9:     Number of leaves at 16 WAP across locations in two planting seasons            62

 4.10:   Number of leaves of some taro genotypes at 20 WAP across two

            locations in 2016 planting season                                                                   63

4.11:    Leaf area of some taro genotypes at 8 WAP across locations in two     

            planting seasons                                                                                              65

 4.12:   Leaf area of some taro genotypes at 12 WAP across locations in two     

            planting seasons                                                                                              66

 4.13:   Leaf area of some taro genotypes at 16 WAP across locations in two     

            planting seasons                                                                                              67

 4.14:   Leaf area of some taro genotypes at 20 WAP across locations in 2016   

            planting season                                                                                               68

 4.15:   Number of secondary suckers of some taro genotypes at 8 WAP across

            locations in 2016 planting seasons                                                                 70

  4.16: Number of secondary suckers of some taro genotypes at 12 WAP across

            locations in 2016 planting seasons                                                                 70

 4.17:   Number of secondary suckers of some taro genotypes at 16 WAP across

            locations in 2017 planting season                                                                   71

 

 4.18:   Number of secondary suckers of some taro genotypes at 20 WAP across

            locations in two planting seasons                                                                   71

 4.19:   Yield (t/ha) of some taro genotypes across two locations in two                        

            planting seasons                                                                                              73

 4.20:   Cormel length (cm) of some taro genotypes across locations in two     

            planting seasons                                                                                              75

 4.21:   Cormel circumference (cm) of some taro genotypes in two locations in     

            2016 and 2017                                                                                                76

 4.22:   Number of cormels per plant of some taro genotypes across two locations

            in two planting seasons                                                                                   77

4.23:    Cormel weight (kg plant-1) of some taro genotype in 2016 planting

            season                                                                                                             80

 4.24:   Cormel weight per plant (kg plant-1) of some taro genotypes in locations          81

 4.25:   Corms length (cm) of some taro genotypes across locations in two                 

            planting seasons                                                                                              85

 4.26:   Corms circumference (cm) of some taro genotypes across two locations in     

            2017 planting season                                                                                      85

 4.27:   Corms weight (kg) per plant of some taro genotypes in two locations as    

            influenced by plant population density                                                          85

 4.28:   Plant height (cm) of some taro genotypes as affected by location and     

            fertilizer in 2016 and 2017 planting seasons at 8 WAP                                 88

 4.29:   Plant height (cm) of some taro genotypes as affected by location and

fertilizer in 2016 and 2017 planting seasons at 12 WAP                               89

 4.30:   Plant height (cm) of some taro genotypes as affected by location and

fertilizer in 2016 and 2017 planting seasons at 16 WAP                               90

 4.31:   Plant height (cm) of some taro genotypes as affected by location and

fertilizer in 2016 and 2017 planting seasons at 20 WAP                               91

 4.32:   Number of leaves at 8 WAP of some taro genotypes as affected by

            locations and fertilizer rates across locations in two years                 94

 4.33:   Number of leaves at 12 WAP of some taro genotypes as affected by

            locations and fertilizer rates across locations in two years                 95

 4.34:   Number of leaves at 16 WAP of some taro genotypes as affected by

            locations and fertilizer rates across locations in two years                 96

 4.35:   Number of leaves at 20 WAP of some taro genotypes as affected by

             locations and fertilizer rates across locations in two years                97

4.36:   Leaf area (cm2) of some taro genotypes at 8 WAP across locations in

two years                                                                                                         100

 4.37:   Leaf area (cm2) of some taro genotypes at 12 WAP across locations in

            two years                                                                                                         101

 4.38:   Leaf area (cm2) of some taro genotypes  at 16 WAP across locations

            in two years                                                                                                    102

 4.39:   Leaf area (cm2) of some taro genotypes at 20 WAP across locations

            in two years                                                                                                    103

 4.40:   Number of secondary suckers (NSS) of some taro genotypes

            at 8 WAP across locations in two years                                             106

 4.41:   Number of secondary suckers (NSS) of some taro genotypes

at 12 WAP across locations in two years                                                       107

 4.42:   Number of secondary suckers (NSS) of some taro genotypes

at 16 WAP across locations in two years                                                       108

 4.43:   Number of secondary suckers (NSS) of some taro genotypes 

            at 20 WAP across locations in two years                                                       109

 4.44:   Yield (t/ha) of some taro genotypes across location in two years                        111

 4.45:   Corm circumference of some taro genotypes across locations in two years         113

 4.46:   Corms length (cm) of some taro genotypes across locations in 2016             114

 4.47:   Cormel length (cm) of some taro genotypes across location in 2017                        114

 4.48:   Corms weight (kg) per plant of some taro genotypes across locations in

                        2017                                                                                                    116

 4.49:   Cormel circumference (cm) of some taro genotypes across location in

            2017                                                                                                                117

 4.50:   Cormel number/plant of some taro genotypes across location in 2017    118

 

4.51:   Plant height (cm) of some taro genotypes at 8WAP across location

            in two years                                                                                                    121

 4.52:   Plant height (cm) of some taro genotypes at 12WAP across location

            in two years                                                                                         122

 4.53:   Plant height (cm) of some taro genotypes at 16WAP across location

in two years                                                                                                    123

 

 4.54:   Plant height (cm) of some taro genotypes at 20WAP across two locations

            in two years                                                                                                    124

 4.55:   Number of leaves of some taro genotypes at 8WAP across locations

in two years                                                                                                    126

 4.56:   Number of leaves of some taro genotypes at 12WAP across locations

            in 2017                                                                                                            127

 4.57:   Number of leaves of some taro genotypes at 16WAP across locations

            in two years                                                                                                    127

 4.58:   Leaf area (cm) of some taro genotypes at 8WAP across

            location in two years                                                                                       129

 4.59:   Leaf area (cm) of some taro genotypes at 12WAP across locations

in 2016                                                                                                            130

 4.60:   Leaf area (cm) of some taro genotypes at 16 WAP across

location in two years                                                                                       131

4.61:    Leaf area (cm) of some taro genotypes at 20WAP across location

in two years                                                                                                     132

 4.62:   Number of secondary suckers (NSS) of some taro genotypes

            at 8WAP across locations in 2017                                                      134

 4.63:   Number of secondary suckers (NSS) of some taro genotypes

at 12WAP across locations in 2016                                                                135

 4.64:  Number of secondary suckers (NSS) of some taro genotypes

at 16WAP across location in two years                                                          136

 4.65:   Number of secondary suckers (NSS) of some taro genotypes

at 20WAP across locations in two years                                               137

 4.66:   Yield (t/ha) of some genotypes at across locations in two years                  139

 4.67:   Corm circumference of some genotypes at across locations in two years    140

 4.68:   Corm length of some genotypes across locations in two years                    142

 4.69:   Cormel circumference (cm) of some genotypes

across location in two years.                                                                           143

 4.70:   Cormel length (cm) of some genotypes across locations in 2016                        144

 4.71:   Number of cormels/plant of some genotypes across locations in two years         146

4.72:    Cormel weight (g) per plant of some genotypes across locations in 2017    148

 4.73:   Corms weight per plant of some genotypes at across locations in 2017    149

4.74:    AMMI analysis of variance for the yield (tha-1) of some taro genotypes

                        grown at 12 environments (combination of 2 locations, two years

            and 3 population density levels)                                                         152

 4.75:   AMMI analysis of variance for the yield (tha-1) of some taro genotypes

                        grown at 16 environments (combination of 2 locations, two years

            and 4 propagule size levels)                                                               161

 4.76:   AMMI analysis of variance for the yield (tha-1) of some taro genotypes       

            grown at 24 environments (combination of 2 locations, two years and 6  

            fertilizer rates)                                                                                                169

 4.77:   Correlation analysis of the growth and yield attributes of taro as

affected by the fertilizer rates in the first planting season (2016)    177

 4.78:   Correlation analysis of the growth and yield attributes of taro as affected

            by the fertilizer rates in the second planting year (2017)                              179

 4.79:   Correlation analysis of the growth and yield attributes of taro as

            affected by the propagule size in the first planting year (2016)                   181

 4.80:   Correlation analysis of the growth and yield attributes of taro as

            affected by the propagule size in the second planting year                            183

 4.81:   Multiple regression (B, step wise) coefficient of determination (R2), R2          

            change (∆R2) between yield (t/ha) and other attributes for some taro     

            genotypes using different fertilizer rates in 2016 and 2017                           185

 4.82:   Multiple regression (B, step wise) coefficient of determination (R2), R2

            change (∆R2) between yield (t/ha) and other attributes for some taro

            genotypes using different propagule sizes in 2016 and 2017                         186

 

 

 

 

 

 

 

LIST OF FIGURES


1:         The AMMI biplot (IPCA1 vs mean) for the yield (t/ha) of 6 taro                        

            genotypes across 12 environments as influenced by population                        

            density                                                                                                            153

 2:        Polygon view of the GGE biplot showing which taro genotype won

            in which environment as influenced by population density                           155

 3:        Ranking genotypes based on both mean and stability relative to an ideal           

            genotype as influenced by population density                                               156

 4:        Ranking environments based on both mean and stability relative to

            an ideal environment as influenced by population density                            157

 5:        The discrimination and representativeness view of the GGE biplot             158

 6:        The AMMI biplot (IPCA1 vs mean) for the yield (t/ha) of 6 taro                        

            genotypes across 16 environments (propagule size-based)                            162

 7:        Polygon view of the GGE biplot showing which taro genotype won

            in which environment (propagule size-based)                                               164

 8:        Ranking genotypes based on both mean and stability relative to an ideal           

            genotype (propagule size-based)                                                                    165

 9:        Ranking environments based on both mean and stability relative to an

            ideal environment (propagule size-based)                                                      166

 10:      The discrimination and representativeness view of the GGE biplot

            to show the discriminating ability and representativeness of the test      

            environments (propagule size-based)                                                            167

 11:      The AMMI biplot (IPCA1 vs mean) for the yield (t/ha) of 6 taro

            genotypes across 24 environments (fertilizer rate-based)                              170

 12:      Polygon view of the GGE biplot showing which taro genotype won

            in which environment (fertilizer rate based)                                                  172

 13:      Ranking genotypes based on both mean and stability relative to an ideal           

            genotype (fertilizer rate based)                                                                       173

 14:      Ranking environments based on both mean and stability relative to

            an ideal environment (fertilizer rate based)                                                    174

 15:      The discrimination and representativeness view of the GGE biplot           

            to show the discriminating ability and representativeness of the test      

            environments (fertilizer rate based)                                                              175

 



 

 

 

 

CHAPTER 1

INTRODUCTION

Cocoyam is the common name for two tuber crops Colocasia esculenta and Xanthosoma sagitifolum, Onyeka (2014). Cocoyam is a stem tuber that is widely cultivated in the tropical regions of the world and is a well known food plant which has a long history of cultivation (Ojiaku et al., 2007).  Cocoyam is the third most important staple root/tuber crop after yam and cassava in Nigeria (Bandyopadhyay et al., 2011). The crop provides a cheaper yam substitute especially during period of food scarcity in many parts of Igbo land (Osawaru and Ogwu, 2014). The Nigerian Academy of Sciences noted that cocoyam may not after all be a “poor man’s food” or “a woman crop” but a crop of promising economic value (FAO, 2009). Nigeria happens to be the largest producer of cocoyam in the world with the production figure of about 1.8 million tonnes per annum (CBN, 2001) and the production is being carried out by rural farmers who employ primitive technology and traditional practices. In fact, Onwueme (1999) had observed that the overall picture of cocoyam in Nigeria is that of a crop that is “casually produced and consumed” while Ezedinma, (1987) had reported that and the totality of published work on cocoyam is insignificant compared to those of rice, cassava and cowpea. Although it has been for centuries the traditional staple in many developing countries, it has received least attention by most national research institutes, extension services and agricultural development planners, despite its nutritional value and industrial uses.

Cocoyam is grown as a root crop because of its edible corms and leaves, which are used as spinach throughout the humid tropics. Okpul (2005) reported that the crop is now accepted as a crop that can guarantee food security, because it is relatively low-priced and could therefore feed many low income families. Nutritionally, cocoyam is composed of 70– 80% water, 20–25% starch and 15–30% protein (Enyinnaya, 1992). Its leaves are excellent sources of folic acid, vitamin C, riboflavin and vitamin A. The flour produced from cocoyam is comparable in properties and bread qualities to that from wheat. Lebot et al. (2006) reported also that cocoyam corms and cormels are rich in mucilage, which can be utilized in the paper industry, in medicinal tablet manufacturing and in traditional medicine practice.

Taro is an important staple crop in rural African countries but its contribution to food security is limited by lack of research on its agronomy and commercialization (Marc, 2009). It has relatively small size starch grains which are easily digestible and therefore acclaimed to be a very good source of carbohydrate for diabetic patients (Vinning, 2003). The corms may be cut up and boiled in curries or fried to make crispy. Leaves and leafstalks can also be cooked and eaten like spinach. It is also used as ornamental in Australia, Japan, Italy and elsewhere. Its ability to tolerate salinity makes it suitable for localities where few other crops grow (Grubben and Denton, 2004).

Although it is an important staple food crop in many tropical countries, especially Nigeria, yield is still low as a result of poor genetic improvement (Ogbonna et al., 2015). There is therefore the need to enhance the production of cocoyam as this will help to increase food security and alleviate poverty among rural people.

A successful cultivation of any crop or cultivar in an agro-climatic location depends on its adaptability and its yield stability. Gene by environment interaction (GEI) normally causes versions in yield performance across environments. The perturbation of GEI is a critical puzzle for plant breeders and unraveling this has brought about more ideas and advances in knowledge of the factors that affect plant boom and improvement (Xu, 2010).

Yield stability refers to a genotype's ability to perform consistently at high or low yield throughout a wide range of environments (Lin and Binns 1994). Assessing the yield and performance of crop sorts across an extensive variety of environments are critical to permit a plant breeder to choose high yielding and continuously performing varieties (Nwangburuka et al., 2011).

Several statistical techniques for reading GEI results were advanced (Eberhrat and Russe, 1966, Kang, 1990; Crossa, 1990; Yan, 2001). Amongst these techniques, additive important outcomes and multiplicative interaction (AMMI) (Gauch, 2006) and Genotype plus Genotype X environment interactions (GGE, biplot) Yan et al., 2007), models are generally used for multi-surroundings trial.

AMMI and GGE biplot analyses are beneficial for easy graphical rationalization of complicated genotype through surroundings and AMMI and GGE make use of important element analysis. GGE biplot differs from AMMI based on how the 2 - way table G X E approach is handled before appearing singular value decomposition (SVD). The AMMI applies SVD to the records minus the genotype and surroundings method, while GGE biplot applies SVD to the information minus the surroundings means simplest (Gauch, 2006). Following, the GGE biplot technique composed of ideas, the biplot idea and GGE concept, these standards are normally used to examine results of web page regression evaluation of data. GGE biplot better suits for mega surroundings evaluation, genotype evaluation (mean vs. stability) and study surroundings evaluation which presents discrimination elasticity or representatives (Amira et al., 2013). The popularity of GGE biplot is related to its versatility and ability to investigate a range of facts with a two-way shape.

Genotype and environment interaction are when two different genotypes respond to environmental variation in different ways. Genotype and environment (G & E) interactions are of major concern to plant breeders. Purely environmental effects, reflecting the different ecological potential of sites and management conditions, are not of direct concern for the breeder or recommendation of plant varieties. Genotypic main effects (i.e. differences in yield between genotypes) provide the only relevant information when genotype x environment (GE) interaction effects are absent or ignored (Eze et al., 2016). However, differences between genotypes may vary widely among environments in the presence of GE interaction effects as large as those reported in extensive investigations (Delacy et al,  1990; Annicchiarico, 1997).

Crop management systems create novel systems that are resilient in the face of changes in climate and rural demographics. Agriculture is constantly adapting to change; consider the revolutions in agriculture due to irrigation; fertilizer; planting density, propagule size, insect, and disease control; and modern tillage systems. We will need to make similar changes to our cropping systems as we face future changes in our climate and an increased need for using resources efficiently (Apantaku, 2000)

The crop responses to planting density varies from species to species and is highly dependent on such environmental conditions as soil characteristics, biotic elements and climatic conditions of the area. Plant spacing according to Baby et al., (2001) involves the growing of plants on a plot of land with sufficient space between each of the plants so that they can develop their roots and shoots fully. Zarate et al., (2007) have reported that corm and cormel yield in taro were highest at a density of 120,000 plants/ha compared to yields obtained at 80,500 and 100,000 plants/ha populations. However, cormel weight decreases with increase in plant population (Osundare, 2007). Orji et al (2014) also have reported that the highest total yield of taro was obtained with close spacing which was contrary to the result obtained by Atiquzzaman et al., (2008). Ukpabi and Nwosu (2010) reported cocoyam yield can be increased by increasing population but without increasing amount of planting material.  Yield can be improved by increasing the individual potentials of each plant or by increasing the yield per surface unit (through high planting densities).

Cocoyam as an important tuber needs nutrients and moisture for vigorous growth (Badaruddin et al., 1999). Soils in this area lack essential nutrients especially those that enhance growth and development in cocoyam.  Fertilizer requirement vary depending on the soil type, native fertility, previous cropping, cultural practices and yield levels (FAO, 2013). The use of inorganic fertilizers in cocoyam production is not common among the rural farmers as they depend more on farmyard manure and household wastes but the quantities of these materials available may not be enough for large scale production. There is then the need to adopt the use of inorganic fertilizers by these farmers. However, one of the problems facing rural farmers on fertilizers usage is lack of information on fertilizer types and quantity.  NPK fertilizers are an all –in –one source of plant nutrients for the individual crops and soils. The use of NPK will provide balanced nutrition and enhance soil fertility, maximize crop yield and ultimately will boost the agricultural economy. Chukwu and Eteng (2014) reported a higher yield of Xanthosoma mafafa with application of 400kg/ha NPK fertilizer in Umudike. Onwueme (1987) reported that cocoyam requires a lot of potassium which in the traditional farming system is found in ash left after bush burning. Information on N.P.K. fertilizer rates of cocoyam and its effects on the genotypes is scanty in literature and hence needs to be worked on. Propagule size is another factor that may affect the growth and yield of taro genotypes. Like most of the tropical root crops, the planting material for taro is bulky, making it expensive to transport over long distances (Mandal et al., 2013). The availability of planting materials is frequently a limiting factor in taro production. With planting materials being so scarce, it is not surprising that some farmers use whatever planting material they can get. Establishing the effect of propagule size on taro may be a solution to reduce the bulkiness of taro planting materials. The use of small set is a means of rapid multiplication of limited quantities of planting materials in the shortest possible time. Smaller sets are planted when the rains have become regular and they are planted 25cm apart on well prepared ridges spaced one metre apart (Ukpabi and Nwosu, 2010). There is limited research work on the size of propagules of taro and research into various agronomic practices is needed in order to improve its productivity and yield stability.

This study was therefore aimed at the following objectives;

1.     To determine yield and yield stability of some taro genotypes at different planting densities.

2.     To determine the effect of N.P.K. fertilizer rates on the yield and yield stability of some taro genotypes.

3.     To determine the yield and yield stability of taro planted with different propagule sizes.

4.     To determine inter-relationship between yield and associated traits of some taro genotypes.

 

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