A STUDY OF FLORISTIC COMPOSITION AND ASSEMBLAGE GUILD OF AVIFAUNA IN COMMUNITY FORESTS OF IKWUANO LOCAL GOVERNMENT AREA, ABIA STATE, NIGERIA

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ABSTRACT

This work was carried out to determine the floristic composition, abundance and assemblage guild of avifauna in community forestof Ikwuano area of Abia State, Nigeria. Twelve (12) plots measuring 60m x 60m each were mapped in 4 major communities of Ikwuano (Oboro, Ibere, Oloko and Ariam). Within each plot, three line transects measuring 20m x 60m were marked to facilitate access to the terrain. The three transects were then divided into twelve subplots, each measuring 10m x 10m for the purpose of identification and complete enumeration of plant species that attains a minimum height of 16cm and above. Enumerated plant species were tagged to avoid repeated counting of individual species and unknown plant species were collected and identified in the herbarium unit of Forestry and Environmental Management Department, Michael Okpara University of Agriculture, Umudike.  A total of eighty nine (89) plant species belonging to forty six (46) families were recorded, of which Dactyladenia barteri had the highest percent relative abundance (8.96%), while Elaeis guineensis and Dialium guineense had 7.01% and 5.68% relative abundance respectively. The following plant families Leguminosae-Pap., Moraceae and Rubiaceae were represented by six (6) plant species, while Apocynaceae, Euphorbiaceae and Malvaceae were represented by four (4) plant species each. Other families were represented by 1 or 3 plant species. Of the 89 plant species recorded, 54 were categorized as Least Concern (LC), 3 Near Threatened (NT), 3 Vulnerable (VU) and 2 Data Deficient (DD) in the IUCN red list of threatened speciesGnetum africanaIrvingia gabonensisand Milicia excelsa were the NT plant species recorded. Also, Terminalia ivorensisNauclea diderrichiiGarcinia sp. were the VU plant species documented in the area. Higher number of plant species diversity were recorded in Ariam, while Oloko had more plant species abundance. Regarding the bird species of the site, aural and visual observation were adopted along the trail of 3 line transects of 20m x 60m of the 60m x 60m main plot with the aid of binoculars for distant birds and mist net for trapping of birds. Photographs of birds caught were taken to validate their occurrence at the site as well as their identification. 25 families comprising of 69 bird species were recorded. Among the documented birds, 46 species were observed within the area, while 23 species were absent during the study but were mentioned in the textual literature referred to. Also, previously unrecorded occurrences of Vidua chalybeataEuplectes hordeaceus and Ardea melanocephala were documented in Ibere, which were absent from the existing literature at the time of this study. Two (2) bird families (Ardeidae and Accipitridae) of National Conservation interest were recorded in the area. The Ibere community exhibits a higher level of bird species richness and diversity compared to Ariam, Oboro and Oloko. The consensus ecological niche modelling used for variable selection showed that Bende, Isialangwa South, Osisioma Ngwa, Ohafia and Arochukwu possess ecological potentials to support similar species assemblages as those documented in Ikwuano. The study’s findings suggest that Ikwuano should be incorporated into the distribution map of newly documented bird species as a recommended course of action. Continuous monitoring exercise to track changes in bird assemblages over time should be adopted so as to provide valuable information on species population trend. To safeguard, enhance and sustain biodiversity stability, it is recommended to engage the community and carry out habitat enhancement activities, particularly in areas that support species categorized as Vulnerable, Near Threatened and of National Conservation interest.







TABLE OF CONTENTS

Title page                                                                                                                    i

Declaration                                                                                                                 ii

Certification                                                                                                               iii

Dedication                                                                                                                  iv

Acknowledgements                                                                                                    v

Table of Content                                                                                                         vi

List of Tables                                                                                                              vii

List of Figures                                                                                                             viii

List of Plates                                                                                                               ix

Abstract                                                                                                                      x

CHAPTER 1:            INTRODUCTION

1.1       Background of the Study                                                                                1

1.2       Statement of Problem                                                                                     3

1.3       Objectives                                                                                                       4

1.4       Significance of the Study                                                                               5

CHAPTER 2:            LITERATURE REVIEW

2.1       Brief Description of Flora                                                                              6

2.1.1    Vegetation                                                                                                      6

2.1.2    Distribution of tropical plants                                                                        7

2.1.2.1 Mangrove forest                                                                                             8

2.1.2.2  Freshwater swamp forest                                                                               9

2.1.2.3  Riparian forest                                                                                               9

2.1.2.4  Lowland rainforest                                                                                        10

2.1.3     Habitat fragmentation                                                                                   12

2.2       Species Diversity                                                                                           12

2.2.1     Plant diversity                                                                                                13

2.3       Species-Area Relationships                                                                           14

2.3.1    The habitat diversity hypothesis                                                                     15

2.3.2    The area per se hypothesis                                                                              16

2.3.3    The passive sampling hypothesis                                                                   16

2.3.4    The resource concentration hypothesis                                                          17

2.3.5    Edge effects                                                                                                   17

2.4       Land Use                                                                                                        18

2.5       Avifauna                                                                                                         20

2.5.1    Classification of avifauna                                                                             22

2.5.2    Life cycle of a bird                                                                                        23

2.5.2.1 The egg stage                                                                                                  23

2.5.2.2 Hatching stage                                                                                                23

2.5.2.3 Nestling stage                                                                                                 23

2.5.2.4 Fledgling stage                                                                                               23

2.5.2.5 Juvenile stage                                                                                                 24

2.5.2.6 Sub adult stage                                                                                                24

2.5.2.7 Adult stage                                                                                                      24

2.5.3    Avifaunal diversity in Nigeria                                                                      24

2.5.4    Bird species assemblage                                                                                25

2.5.4.1 Flock                                                                                                               25

2.5.4.2 Colony                                                                                                            26

2.5.4.3 Mixed-species foraging flock                                                                         26

2.5.4.4 Swarming                                                                                                        26

CHAPTER 3:            MATERIALS AND METHOD

3.1       Study Area                                                                                                      21

3.2.      Plant species data collection                                                                          30

3.3       Bird Species Data Collection                                                                         32

3.4       Species Distribution Map                                                                               33

3.5       Data Analyses                                                                                                 34

3.5.1    Environmental variables and ecological niche modelling                             34

 

CHAPTER 4:            RESULTS AND DISCUSSION

4.1       Results                                                                                                            36

4.1.1    Plant species composition                                                                              36

4.1.2    Percent frequency of occurrence of plant species in the site                          40

4.1.3    List of IUCN status of plant species in the site                                              43

4.1.4    Plant habit, number of generic and species composition in the site                        45

4.1.5    Bird species composition                                                                               46

4.1.6    Bird species caught and photographed during the study                                50

4.1.6.1 Bronze Mannikin (Spermestes cucullatus)                                                     50

4.1.6.2 Adult Female of Yellow-Mantled Widowbird (Euplectes macroura)       52

4.1.6.3 Blue-Spotted Wood Dove (Turtur afer)                                                         53

4.1.6.4 Simple Leaflove (Chlorocichla simplex)                                                       54

4.1.6.5 Yellow-Rumped Tinkerbird (Pogoniulus bilineatus)                                    55

4.1.6.6 Grey-Backed Camaroptera (Camaroptera brachyura)                                  56

4.1.6.7 Magpie Mannikin (Spermestes fringilloides)                                                 57

4.1.6.8 Black-Winged (Red) Bishop (Euplectes hordeaceus)                                    58

4.2       Discussion                                                                                                       59

4.2.1    The status of recorded plant species on IUCN red list of                              

threatened species                                                                                           60

4.2.2    Bird species record and their status on IUCN red list of

threatened species                                                                                           60

4.2.3    Ecological niche modelling                                                                            67

CHAPTER 5:     CONCLUSION AND RECOMMENDATION

5.1       Conclusion                                                                                                      70

5.2       Recommendation                                                                                           71

 

REFERENCE                                                                                                            73

APPENDICES                                                                                                           84


 






LIST OF TABLES

 

4.1:      Checklist of Plant Species Composition and their Percentage Relative

Abundance in the Study Sites                                                             37

4.2.      Percentage Frequency of Occurrence of Plant Species in the Study Site       41

4.3.      Checklist of IUCN Status of Plant Species in the Study Site                         43

4.4.      Plant Habit, Number of Generic and Species Composition                           46

4.5       Checklist of Bird species documented at the site and their IUCN Status  47

4.6.      Bird species richness, diversity, and evenness across the survey areas    62









 

LIST OF FIGURES

 

 

1.       The rainforest canopy (Source: Butler, 2006).                                             11

 

2.       The Species–area curve of vascular plants. The logarithm of species

richness is commonly plotted as a function of the logarithm of area.

(Source: Connor and McCoy, 1979).                                                             14

 

3.         Relative effect of major drivers of biodiversity change.

(Source: Sala et al., 2000).                                                                             18

 

4:        Framework of potential transitions between land-use/cover states.

(Source: Lambin and Geist, 2006)                                                                  19

 

5:        A diagram of Bird (Source:  Garrigues and Dean, 2014)                             21

 

6.         Geologic map of Abia State showing Ikwuano

(Source: Amos-Uhegbu et al., 2014)                                                              28

 

7.         Map showing the bordering communities of the site

(Source: Amos-Uhegbu et al., 2014)                                                              29

 

8.         Diagram of field plot layout showing 60m x 60m main plots,

20m x 60m transect plots containing 10m x 10m subplots.                           31

 

9.         The differences in bird species assemblage in surveyed locations

of Ikwuano LGA                                                                                             63

 

10.       The differences in bird species identified based on the different

observation methods in surveyed locations of Ikwuano LGA.                        64

 

11.       The relationship between bird species richness and plant species

abundance in each surveyed location of Ikwuano LGA.                                65

 

12.       The relationship between bird species richness and plant species

richness in each surveyed location of Ikwuano LGA.                        66

 

13.       The distribution of potential habitats with similar to the surveyed

location of Ikwuano LGA based on the svme Ecological

Niche Prediction algorithm.                                                               68

 

14.       The distribution of potential habitats with similar to the

surveyed location of Ikwuano LGA based on the consensus

model of Ecological Niche Prediction algorithms.                            69


 







LIST OF PLATES

 

3.1: A mist net set for the trapping of avifauna species                                             33

4.1: Juvenile of Bronze Mannikin (Spermestes cucullatus)                                       51

4.2: Adult Female of Yellow-Mantled Widowbird (Euplectes macroura)                        52

4.3: Blue-Spotted Wood Dove (Turtur afer) entrapped in Mist net                           53

4.4: Simple Leaflove (Chlorocichla simplex)                                                            54

4.5: Yellow-Rumped Tinkerbird (Pogoniulus bilineatus)                                         55

4.6: Grey-Backed Camaroptera (Camaroptera brachyura)                                       56

4.7: Magpie Mannikin (Spermestes fringilloides)                                                      57

4.8: Black-Winged (Red) Bishop (Euplectes hordeaceus)                                        58


 





 

 CHAPTER 1

INTRODUCTION

1.1 BACKGROUND OF THE STUDY

Multitudes of different species of living organisms are facing much coercion and, as a result, the disappearance of wild plant and animal diversities is continuously increasing and taking place at an appalling short-time scale (Novacek and Cleland, 2001). Although, the actual total diversity of biological species on earth is not yet known (May, 1992). According to Attiwill (1994) changes in conditions created by disturbances generate structural and compositional diversities in the forest ecosystem. The effects of these threats may be insignificant until the affected species population begin to experience conspicuous decline. According to Pimm et al., 1995 and Brooks et al., 2006, anthropogenic actions are causing biodiversity crisis, with species extinction rating up to 1000 times higher than background. Habitat destruction has been reported as one of the major threats to biodiversity (Myers et al., 2000; Thomas et al., 2004), and the diversity of plants, bird and insect species successfully thrive in the forest ecosystems (Nadkarni, 1994).

The extent of forest habitat as reported by Raf and Olivier (2011) has continued to decrease due to forest conversion to agriculture, and land uses associated with urban population growth. Between 1980 and 2000 more than 50 percent of the recent agricultural lands across the tropics were acquired by clearing intact forests (Raf and Olivier, 2011). The population pressure has unleashed some notable threats to ecosystems and biodiversity, which include overexploitation of forest resources, invasion of alien species, loss of habitat and fragmentation, pollution, global climate change and disruption of community structures (Chape et al., 2003; Novacek and Cleland, 2001).

Plant species are one of the major ecological components of forest. They form the physical structure which comprises of tall and short plants occupying space in various configurations of tightness or openness (Patton, 2010). Plant species generally form the vegetation types of habitat and ecosystem at large. They are known to be the major primary producer in the food chain, and have other consumers that depend on them for food. There are indicator plant species used in marking zones as classified by Keay (1959). Those species are most recurrently encountered in the natural vegetation of the zones (Isichei, 2015). Plant communities have a substantial influence on the distribution and interaction of animal species, as they determine the physical structure of the environment which supports the guild of birds, mammals and other animal species within the forest (Lawton, 1983). So many authors such as MacArthur et al., (1962), MacArthur, (1964) and Joshi et al., (2012) have argued on the determinant of bird species distribution. They reported that, in determining the bird species diversity, vertical diversity of foliage may be more crucial than the actual composition of plant species. Bird and habitat relationship according to Emien (1977), Verner et al., (1986) and Jones et al., (1996) may also be determined by vegetation physical-structural variables such as canopy cover, sizes and height of trees etc.

More so, Wiens and Rotenberry (1981) reported that locations of higher habitat diversity have the tendency of supporting the activities of most bird species. Biodiversity assessment of various habitats has been encouraged and extensively adopted as one of the crucial steps towards effective management of the environment and its related threats (Olson, et al., 2001; Oliver and Beattie, 1993). One of the ideal productive approaches for conserving the variety of life on earth is by identifying places of high biological diversity (i.e. biological hotspots) and subsequently developing and implementing management strategies for sustaining and improving the conservation status of such areas (Ezealor, 2002). Consequently, this study meant to identify the active areas where plant and bird species under threat are found in Ikwuano, and attention called for their protection.


1.2 STATEMENT OF PROBLEM

The coverage of forest habitats which are homes to thousands of plant and animal species has continued to decrease due to unstoppable conversion of naturally existing forested areas to agricultural farmlands, and residence for urban population growth by humans (Raf and Olivier, 2011). This has led to the displacement of the natural inhabitants of the forest apposite to play important roles in the biosphere transfer of energy and nutrients.

According to Obi (2002), the loss of biological diversity in Nigeria has become a serious issue due to their importance in human’s economic and social activities. Despite the importance of biodiversity, humans have caused a near destruction of the ecosystem which remains a serious threat to development and sustainable living (Obi, 2002). When the quality and integrity of a habitat is compromised, the inhabitant species of the site become vulnerable to destruction (Gurevitch and Padilla, 2004). Habitat loss is a major threat to biodiversity species (Henle et al., 2008), and series of efforts towards its conservation has continued to progress. A series of conservation approaches have been adopted to decrease the rate of loss. According to Ezealor, 2002 the tradition and practices of various cultures in Nigeria revolve around protection of habitats and species.

Long before now, some communities conserved forests within their localities with intent for hunting exercise while others established sacred groves for their idol praise. In some cases, totem and taboos were used to conserve individual plant and animal species valued for the purposes of medicine, shade and food. National parks, game reserves and forest reserves are the organised conservation strategies in Nigeria and in some other countries. According to Ogunjinmi et al., (2017), the aspiration of a nation to preserve the flora and fauna as elements of its national and cultural heritage for the generations unborn is reflected in the establishment of parks. Human habitation and other activities that affect biodiversity are disallowed in the conservation area. However, with increase in population pressure and lack of interest and commitment, these measures are challenged. Consequently, Ezealor (2002) reported an approach for biodiversity conservation using bird species as a tool in Nigeria. According to International Council for Bird Preservation/BirdLife International (ICBP) (1992) birds are good indicators of the health of the environment and as well used to predict the distribution and conservation status of other types of animals and plants.

The paucity of literature on the locations that accommodate threatened plant and bird species in Ikwuano triggered the quest for this research. This could be as a result of habitat destruction which may detract survey attention. Though, patches of isolated forests still exist. This study will help identify and document areas of plant and bird species of global interest in order to facilitate their protection and to avoid total loss of the species.


1.3       OBJECTIVES

The aim of the research is to study the flora composition and assemblage guild of avian species in community forests of Ikwuano. The specific objectives included;

i)               to provide a check list of the flora and avifauna species of Ikwuano,

ii)             to determine their current global threat status, and

iii)           to produce species distribution map of locations with related flora and avifauna composition of the study site in Abia State using ecological niche modelling.


1.4       SIGNIFICANCE OF THE STUDY

Tropical rainforests due to their evergreen vegetative makeup harbour diversity of species, but deforestation and over-exploitation encroachments have to some extent reduced their potentials to serve the purpose of biodiversity reservoirs (Geist and Lambin, 2002). According to Pereira et al., (2010) the decline of both plant and animal species in the tropical rainforests and other ecosystems has become one of the major environmental concerns of 21st Century. However, monitoring and documentation of the existence and activities of biodiversity is vital to its conservation (Singh, 2002).

The assessment of biodiversity in an ecosystem is important in the design and formation of adequate management plan (Richardson, 1999; Verma et al., 2004). Having been considered a good ecological practice as stated by Onyenweaku et al., (2018) many habitats and species have been under regular check. This assessment ensures recording of species which could make known the biotic integrity of the ecosystem (Karr, 1981). Some scientific authors such as Bibby and Burgess (1992), Roy et al., (2012) reported that ornithological stock-taking is the most common and simplest practice used to describe where birds are found. This approach is ideal because birds play various essential ecological roles in the ecosystem including pollination of plant species, dispersal of seeds of some plant species, bio-indicator etc. (Kumbar and Ghadage, 2014). Most tropical forest habitats serve as destination points for migratory birds (Sanderson et al., 2006). The locations to which these migratory species migrate to, need to be identified and efforts made towards ensuring that the structures which may be the attractant for them are not destroyed.

 

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