PRODUCTION OF ETHANOL FROM CASSAVA PEELS USING ZYMOMONAS MOBILIS AND SACCHAROMYCES CEREVISIAE

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ABSTRACT

The yield of bioethanol from cassava peels hydrolysed with Aspergillus niger and crude enzymes (amylase, cellulase and pectinase) was examined in this study. Cassava peels were pretreated by soaking and a combination of soaking and boiling which removes 67% of cyanide. After hydrolysis, Zymomonas mobilis and Saccharomyces cerevisiae were used to ferment the hydrolysates separately at room temperature for five (5) days. The result showed that pretreatment by soaking and boiling for up to 120 minutes removed the highest amount of cyanide and increased carbohydrate but reduced the fibre content (37.04±0.01mg/g, 71.42±0.02%, and 10.38±0.42%). Hydrolysis using Aspergillus niger yielded up to 95.44±0.11mg/g reducing sugar while hydrolysis using enzymes yielded up to 72.38±0.06mg/g reducing sugar.  Highest ethanol yield when the hydrolysates were fermented with Zymomonas mobilis and Sacchromyces cerevisiae were 3% and 2% respectively. Proximate composition of the fermented waste showed 6.37±0.08, 2.71±0.03, 25.49±0.01, 8.97±0.02 and 34.41±0.16% ash, fat, protein, crude fibre and carbohydrate respectively while nitrogen, phosphorus and potassium of the residue was 4.08±0.50, 0.40±0.28 and1.84±0.15% respectively. The work reveals that cassava peels is a suitable agricultural waste for bioethanol production and fermentation with Zymomonas mobilis yielded high ethanol compare to Saccharomyces cerevisiae. The work also proved that the waste product of fermentation may be used as animal feedstock or fertilizer to enrich the soil for plant growth.    

 

 

 

 

 

 

 

 

 

 

 

 

TABLE OF CONTENT

Title page                                                                                                                                i

Declaration                                                                                                                             ii

Dedication                                                                                                                              iii

Acknowledgement                                                                                                                  iv

Abstract                                                                                                                                   v

Table of content                                                                                                                      vi

CHAPTER ONE                                                                                                                     1

INTRODUCTION                                                                                                                  1

1.1 QUEST FOR ALTERNATIVE ENERGY SOURCE                                                      2

1.2 JUSTIFICATION                                                                                                             2

1.3 STATEMENT OF RESEARCH PROBLEM                                                                   2

1.4 RESEARCH AIM AND OBJECTIVES                                                                          2

CHAPTER TWO                                                                                                                    4

LITERATURE REVIEW                                                                                                       4

2.1 CASSAVA PRODUCTION IN AFRICA                                                                        4

2.2 CASSAVA PEELS                                                                                                           4

2.3 CYANIDE IN CASSAVA                                                                                                5

2.4 LIGNOCELLULOSIC BIOMASS                                                                                  6

2.4.1 SECOND GENERATION FEEDSTOCKS                                                                 7

2.4.2 STRUCTURE OF LIGNOCELLULOSIC BIOMASS                                                8

2.5 STRUCTURE AND PROPERTIES OF STARCH                                                          12

2.6 PRETREATMENT OF LIGNOCELLULOSIC BIOMASS                                            14

2.6.1 MECHANICAL PRETREATMENT                                                                           16

2.6.2 PHYSICAL PRETREATMENT                                                                                   16

2.6.3 PHYSICOCHEMICAL PRETREATMENT                                                                 17

2.6.4 CHEMICAL PRETREATMENT                                                                                  17

2.6.5 BIOLOGICAL PRETREATMENT                                                                              18

2.7 HYDROLYSIS AND FERMENTATION OF LIGNOCELLULOSIC BIOMASS       19

2.8 BIOETHANOL                                                                                                                 20

2.8.1 GLOBAL BIOETHANOL PRODUCTION                                                                  21

2.8.2 PRODUCTION AND UTILIZATION OF ETHANOL INNIGERIA                22

2.9 MICROORGANISMS EXPLORED FOR ETHANOL PRODUCTION                                   22

2.9.1 MICROORGANISMS OF INTEREST IN THIS RESEARCH WORK                    23

2.9.2 Aspergillus niger                                                                                         23

2.9.3 Zymomonas mobilis                                                                                                       24

2.9.4 THE METABOLIC PATHWAY OF Zymomonas mobilis                                           25

2.9.5 Saccharomyces cerevisiae                                                                                             27

2.10 MECHANISM OF YEAST FERMENTATION                                                            28

2.10.1 INHIBITION OF ETHANOL PRODUCTIVITY IN YEAST CELLS                        29

2.11 HYDROLYTIC ENZYMES                                                                                          30

2.11.1 MEASUREMENT OF ENZYME ACTIVITY                                                           31

2.11.2 AMYLASE                                                                                                                  31

2.11.3 CELLULASE                                                                                                               32

2.11.4 PECTINASE                                                                                                                33

2.12 FERMENTED CASSAVA PEELS AS ANIMAL FOOD SUPPLEMENTS                    34

CHAPTER THREE                                                                                                                35

MATERIALS AND METHODS                                                                                           35

3.1 Collection of sample (Cassava peels)                                                                               35

3.2 Microorganisms                                                                                                                35

3.3 Identification of microorganisms                                                                                     35

3.3.1 Aspergillus niger                                                                                                           35

3.3.2 Saccharomyces cerevisiae                                                                                             35

3.4 Isolation of Zymomonas mobilis from fresh sweet Palm Wine                                        36

3.5 Pretreatment of Cassava peels                                                                                          36

3.6 Determination of cyanide in pretreated cassava peels                                                      36

3.7 Determination of crude fibre in pretreated cassava peels                                                 37

3.8 Determination of Carbohydrate                                                                                        37

3.9 PRODUCTION OF ENZYMES (Amylase, Cellulase, and Pectinase)                                    37

3.10 Assay for Cellulase activity                                                                                           38

3.11Assay for pectinase activity                                                                                            38

3.12 Assay for amylase activity                                                                                             39

3.13 Enzyme Hydrolysis                                                                                                        39

3.14 Microbial Hydrolysis                                                                                                      39

3.15 Determination of Reducing Sugar                                                                                  40

3.16 Fermentation of the Hydrolysate                                                                                    40

3.17 Distillation                                                                                                                      40

3.18 Determination of Percentage Ethanol Yield                                                                   40

3.19 PROXIMATE ANALYSIS OF THE HYDROLYSATE                                               41

3.19.1 Total ash                                                                                                                      41

3.19.2 Crude fat                                                                                                                      41

3.19.3 Crude fibre                                                                                                                   41

3.19.4 Crude Protein                                                                                                               42

3.20 Determination of Phosphorus in the Waste after fermentation                                      42

3.21 Determination of Nitrogen in the Waste after fermentation                                          42

3.22 Determination of potassium                                                                                           43

3.23 Analysis of data                                                                                                              43

CHAPTER FOUR                                                                                                                  44

4.1 Result                                                                                                                                44

4.2 Discussion of result                                                                                                          52

CHAPTER FIVE                                                                                                                    56

CONCLUSION                                                                                                                       56

RECOMMENDATION                                                                                                          56

REFERENCE                                                                                                                         57

APPENDIX                                                                                                                             77

 

 

 

 

 

 

 

LIST OF TABLES

Table 2.1: Cyanogenic glycosides (CNp) in different tissues of the cassava plant               6

Table 2.2: Composition of some common sources of biomass                                              12

Table 4.1 Morphological and biochemical characterization of isolates from palm wine          44

Table 4.2: Assay for activities of enzymes produced from Aspergillus niger                46

 

 

 

  

 

 

LIST OF FIGURES


Figure 2.1: Location of cyanogenic glycoside and the enzyme linamarase in the cell        5

Figure 2.2: Hydrolysis of linamarin, in β-glucosidase, and in α-hydroxynitrile lyase    6

Figure 2.3: structure of plant cell wall components                                                               8

Figure 2.4: Structure of lignin                                                                                                9

Figure 2.5: Structure of Hemicellulose                                                                                  10

Figure 2.6: Structure of cellulose                                                                                           11

Figure2.7: The structure of Amylose                                                                                     13

Figure2.8: Structure of Amylopectin                                                                                     13

Figure2.9: Schematic representation on biomass pretreatment                                             15

Figure 2.10: The Entner-Doudoroff (ED) pathway and ethanologenesis                                    26

Figure 2.11: Metabolic pathway for ethanolic fermentation in S. cerevisiae cells                       27

Figure 2.12: Possible target sites for ethanol inhibition in yeast cells                                     29

Figure 4.1: The carbohydrate, crude fibre and cyanide contents of the pretreated cassava peel           45

Figure 4.2: Reducing sugar yield from hydrolysis of treated cassava peels using A. niger 47

Figure 4.3: Reducing sugar yield from enzymatic hydrolysis of treated cassava peels   48

Figure 4.4: Ethanol yield by Z. mobilis and S. cerevisiae fermentation                                     49

Figure 4.5: Proximate analysis of dried waste product of fermentation                                    50

Figure 4.6: Nitrogen, Potassium and phosphorus in the waste product of fermentation   51

 

 

           

 

 

CHAPTER ONE

INTRODUCTION

Cassava is one of the important food crops that survive conditions of low nutrient availability and drought (John, 2004). History revealed that cassava originates from Brazil and was introduced into West Africa by Portuguese and today, over 600 million people from Africa, Asia and Latin America depend on cassava for food (Asegbeloyin and Onyimonyi, 2007). Cassava was introduced into Nigeria 300 years ago and became generally accepted and cultivated in late 1990s (Nigeria Cassava Master Plan, 2006). Today, Nigeria is the highest producer of cassava in the world, producing higher than Brazil, Thailand and Indonesia (Asegbeloyin and Onyimonyi, 2007). Cassava has been a raw material globally for industrial production of textiles, papers, adhesives, pharmaceuticals and various food products because it is rich in carbohydrate with high energy density and has generated great impact in world economics (Aigbe and Remison, 2010). Cassava and it is by – products are renewable source of energy that pose no threat to the environment and has been used by several researchers in the production of biofuel (Musatto and Teixira, 2010). Interest in the production of biofuel from agricultural wastes is driven by several reasons such as: global search for alternative source of energy and transportation fuel to replace the depleting fossil fuel. It was also because of several benefits derivable from the use of ethanol. These benefits includes: the use of ethanol as solvents, germicides, antifreeze and intermediates for other organic chemicals (Kingsley, 2012). Bioethanol can be produced by fermenting sugars in plant materials as opposed to synthetic ethanol production from petrochemical sources (Oyeleke a Jibrin, 2009). Ethanol is pollution free, biodegradable, renewable, cause no climate change and the by – product of fermentation can be used as animal feedstock (Adelekan, 2010; Teerapatr, Lerdluk and La-aied, 2010). It has been estimated that 2.96 million metric tons of cassava peels is generated annually in Nigeria from processing cassava to various food products (Nwabueze and Otunwa, 2006). These enormous wastes that constitute 20 – 35% of the weight of cassava tuber are discarded with consequent implication of environmental pollution. In view of this, it has become necessary to convert this waste to useful end products in other not to pose threat to the environment (Nwabueze and Otunwa, 2006).

 

1.1  QUEST FOR ALTERNATIVE ENERGY SOURCES

The earth experienced an increase in the mean temperature in the 19th century due to emission of greenhouse gases. Carbon dioxide has been the largest greenhouse gas emitted through combustion of fossil fuel as coal, oil and natural gases (Sun and Cheng, 2002). United States alone is responsible for 25% of global energy consumption and 25% of the world carbon dioxide emission. Researches have shown that the deposit of crude oil in the earth crust is gradually depleting with time. This reason couple with the continuous hike in the pump price has driven global attention to the search for a renewable energy source to serve as transportation fuel and power industrial machines (Jin, Michel, Wyman and Dale, 2003).  Bioethanol produced from fermentation of sugars in plants has been discovered to be a perfect replacement for gasoline in some advanced countries (Sharma, Kalra, Oberoi and Bansal, 2007).


1.2  STATEMENT OF RESEARCH PROBLEM

 

Large tons of cassava wastes are discarded annually in Nigeria when cassava is processed to various food products. This waste ends up in open dumps or drainage systems, threatening both surface water and ground water quality. It is therefore necessary to convert these wastes to useful end products rather than allowing them to become nuisance to the environment.

 

1.3  JUSTIFICATION

 

The global search for alternative renewable transportation fuel to replace the depleting fossil fuel is on the increase due to hike in petroleum price and global warming. This research work is aimed at investigating the possibility of transforming cassava peels to valuable products like ethanol, animal feedstock or fertilizer, thereby contributing toward alternative energy supply as well as creating employment opportunity.

 

1.4  RESEARCH AIM AND OBJECTIVES

 

AIM

 

The aim of this research work is to produce ethanol from Cassava peels using Zymomonas mobilis and Saccharomyces cerevisiae and compare ethanol yield of the two organisms.

 

 

OBJECTIVES

 

The overall objectives of this work are:

1. To determine the pretreatment method for cyanide removal in cassava peels.

2. To determine the condition for enzyme hydrolysis of pretreated cassava peels.

3. To establish fermentation condition for hydrolyzed cassava peel.

4. To determine the yield of ethanol produced from fermentation by Zymomonas mobilis and Saccharomyces cerevisiae

5. To assess the fermentation residue for possible organic fertilizer or animal feedstock.

 

 

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